Why Do Some U.S. Bills Have a Star at the End of the Serial Number?

dollar bill — This U.S. $1 bill has been marked as a replacement note. Notice the star located at the end of the serial number. BANKNOTEWORLD/HOWSTUFFWORKS

All modern United States currency contains either a 10- or 11-digit serial number in order to make each bill unique. Ten-digit serial numbers were on all bills until the "new style" came out in 1996. Those bills (and all produced since then) have an 11-digit serial. The serial number consists of the following:

The first letter, only found on the new-style bills, represents the series of the bill. The series indicates the year in which the design of the bill was approved for production. This begins with A, and moves through the alphabet each time a new series is needed (for example, each time there is a new secretary of the treasury, the bill design changes because the secretary's signature is on all currency). You can also find the series of the bill printed directly to the bottom-right of the portrait.
The second letter (or first, if you're looking at an old-style bill) represents the district of the Federal Reserve Bank that your bill was issued from. As there are 12 Federal Reserve Banks, this letter can range from A to L, with A representing Boston and L representing San Francisco.
The eight numerical digits that follow represent a unique ID number. This number increases sequentially as each bill is printed. Using these digits alone, there would be a possible 99,999,999 bills issued per bank.
The final letter is used to raise the number of possible bills beyond 99,999,999. Altogether, there are a possible 2,499,999,975 serial numbers for each bank.

On occasion, a bill shows up that has a small star in place of the final letter in the serial, and many early issues carried the star in front of the serial number.

This star represents what is known as a replacement note. When a printing error occurs during a normal press run and renders a set of bills unusable, replacement notes are used instead. With replacements notes, a set of serial numbers can still have the proper number of bills even if some of the original bills had to be pulled. The replacement notes have a sequence of their own, using the star as their final "letter." This allows for 99,999,999 possible replacement notes for any given bank, series and denomination.

In general, replacement notes aren't worth more than regular bills. (It's a different story for printing errors.) However, if you find a replacement note with a particularly interesting serial number — like 00000001 or 999999999 — or a large number of consecutively numbered replacement notes that you keep together as a lot, you may have a collector's item on your hands.

Now That's Interesting

According to Economics Stack Exchange, a site for those who study, teach and research economic issues: "The U.S. Bureau of Engraving and Printing creates all of the nation's bills, while the U.S. mint creates its coins. But they also destroy money. Banks and individuals will hand over 'mutilated' bills and coins to these agencies. They then validate its authenticity and issue a Treasury check in return. The Bureau of Engraving and Printing receives around 25,000 mutilated currency redemption claims annually. Each bill is shredded and sent to waste energy facilities for disposal."